Donald Steven read zoology at King’s College, Cambridge, and was subsequently appointed demonstrator in Veterinary Anatomy there (1961). A talented artist, he investigated the vascular anatomy of the placenta in the sheep and mare, guided by Robert Comline in the Physiological Laboratory, and edited Comparative placentation: essays in structure and function (1975). He subsequently became Director of the Sub-department of Veterinary Anatomy (1984-9). As a Fellow of Churchill College, Cambridge, he was the Physiological Society’s first archivist when its archive was established there in the 1970s.

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